Entity Central Index Key Cover [Abstract] Document Type Document Type Amendment Flag Amendment Flag Document Period End Date Document Period End Date Document Fiscal Year Focus Document Fiscal Year Focus Document Fiscal Period Focus Document Fiscal Period Focus Trading Symbol Trading Symbol Entity Registrant Name Entity Registrant Name Security Exchange Name Security Exchange Name Entity Central Index Key Entity Current Reporting Status Entity Current Reporting Status Current Fiscal Year End Date Current Fiscal Year End Date Entity Filer Category Entity Filer Category Entity Shell Company Entity Shell Company Entity Small Business Entity Small Business Entity Emerging Growth Company Entity Emerging Growth Company Title of 12(b) Security Security12b Title Entity Common Stock, Shares Outstanding Entity Common Stock Shares Outstanding Entity Address, State or Province Entity Address State Or Province Entity File Number Entity File Number Entity Incorporation, State or Country Code Entity Incorporation State Country Code Entity Tax Identification Number Entity Tax Identification Number Entity Address, Address Line One Entity Address Address Line1 Entity Address, Address Line Two Entity Address Address Line2 Entity Address, City or Town Entity Address City Or Town Entity Address, Postal Zip Code Entity Address Postal Zip Code City Area Code City Area Code Local Phone Number Local Phone Number Entity Interactive Data Current Entity Interactive Data Current Document Quarterly Report Document Quarterly Report Document Transition Report Document Transition Report Prepaid research and development noncurrent. Statement Of Financial Position [Abstract] Assets Assets [Abstract] Current assets: Assets Current [Abstract] Cash and cash equivalents Cash And Cash Equivalents At Carrying Value Short-term marketable securities Available For Sale Securities Debt Securities Current Accounts receivable Accounts Receivable Net Current Prepaid expenses and other current assets Prepaid Expense And Other Assets Current Total current assets Assets Current Property and equipment, net Property Plant And Equipment Net Right of use asset - operating lease Operating Lease Right Of Use Asset Restricted cash Restricted Cash Noncurrent Prepaid research and development, net of current portion Prepaid Research And Development Noncurrent Other assets Other Assets Noncurrent Total assets Assets Liabilities and Stockholders' Equity Liabilities And Stockholders Equity [Abstract] Current liabilities: Liabilities Current [Abstract] Accounts payable Accounts Payable Current Accrued expenses Accrued Liabilities Current Deferred revenue Contract With Customer Liability Current Lease liability - operating lease Operating Lease Liability Current Finance lease obligations Finance Lease Liability Current Total current liabilities Liabilities Current Long-term liabilities: Liabilities Noncurrent [Abstract] Lease liability - operating lease, net of current portion Operating Lease Liability Noncurrent Finance lease obligations, net of current portion Finance Lease Liability Noncurrent Other long-term liabilities Other Liabilities Noncurrent Total long-term liabilities Liabilities Noncurrent Commitments and contingencies (Note 11) Commitments And Contingencies Stockholders' equity Stockholders Equity [Abstract] Common stock, $0.001 par value 250,000,000 shares authorized as of June 30, 2021 and December 31, 2020. 52,375,344 shares issued and outstanding as of June 30, 2021 and 38,183,273 shares issued and outstanding as of December 31, 2020. Common Stock Value Additional paid-in capital Additional Paid In Capital Common Stock Accumulated other comprehensive income Accumulated Other Comprehensive Income Loss Net Of Tax Accumulated deficit Retained Earnings Accumulated Deficit Total stockholders' equity Stockholders Equity Total liabilities and stockholders' equity Liabilities And Stockholders Equity Common stock, par value Common Stock Par Or Stated Value Per Share Common stock, authorized Common Stock Shares Authorized Common stock, Issued Common Stock Shares Issued Common stock, outstanding Common Stock Shares Outstanding Income Statement [Abstract] Revenue Revenues Operating expenses: Operating Expenses [Abstract] Research and development Research And Development Expense General and administrative General And Administrative Expense Total operating expenses Operating Expenses Loss from operations Operating Income Loss Other income (expense): Nonoperating Income Expense [Abstract] Interest and investment income Investment Income Nonoperating Interest expense Interest Expense Other expense Other Nonoperating Income Expense Other income (expense), net Nonoperating Income Expense Net loss Net Income Loss Net loss per share - basic and diluted Earnings Per Share Basic And Diluted Weighted-average common stock outstanding - basic and diluted Weighted Average Number Of Share Outstanding Basic And Diluted Comprehensive loss: Comprehensive Income Net Of Tax [Abstract] Net loss Net unrealized (loss) gain on marketable securities Other Comprehensive Income Unrealized Holding Gain Loss On Securities Arising During Period Net Of Tax Comprehensive loss Comprehensive Income Net Of Tax Statement Of Stockholders Equity [Abstract] Statement [Table] Statement [Table] Equity Components Statement Equity Components [Axis] Equity Component Equity Component [Domain] Common Stock $0.001 Par Value Common Stock [Member] Additional Paid-in Capital Additional Paid In Capital [Member] Accumulated Other Comprehensive Income (Loss) Accumulated Other Comprehensive Income [Member] Accumulated Deficit Retained Earnings [Member] Sale of Stock Subsidiary Sale Of Stock [Axis] Sale of Stock Sale Of Stock Name Of Transaction [Domain] At-the-market offering, At-the-market Offering At The Market Offering [Member] Statement [Line Items] Statement [Line Items] Balance Balance (in Shares) Shares Outstanding Proceeds from issuance of common stock, net of issuance costs Stock Issued During Period Value New Issues Proceeds from issuance of common stock, net of issuance costs, Shares Stock Issued During Period Shares New Issues Issuance of common stock under employee stock purchase plan Stock Issued During Period Value Employee Stock Purchase Plan Issuance of common stock under employee stock purchase plan, Shares Stock Issued During Period Shares Employee Stock Purchase Plans Exercise of options Stock Issued During Period Value Stock Options Exercised Exercise of options, shares Stock Issued During Period Shares Stock Options Exercised Issuance of restricted stock Stock Issued During Period Value Restricted Stock Award Gross Issuance of restricted stock, shares Stock Issued During Period Shares Restricted Stock Award Gross Restricted stock awards withheld for payment of employees' withholding tax liability Restricted Stock Value Shares Issued Net Of Tax Withholdings Restricted stock awards withheld for payment of employees' withholding tax liability, Shares Restricted Stock Shares Issued Net Of Shares For Tax Withholdings Cancellation of restricted stock Stock Issued During Period Shares Restricted Stock Award Forfeited Equity-based compensation expense Adjustments To Additional Paid In Capital Sharebased Compensation Requisite Service Period Recognition Value Unrealized gain (loss) on securities Net loss Balance Balance (in Shares) Reduction in carrying amount of operating lease right-of-use asset. Increase (decrease) in prepaid research and development net of current portion. Proceeds from employee stock purchases and exercise of stock options. Assets acquired under operating lease obligation. Noncash purchase of property, plant and equipment in accounts payable and accrued liabilities adjustments. Issuance costs included in accounts payable and accrued expenses. Noncash investing items. Noncash financing items. Statement Of Cash Flows [Abstract] Cash flows from operating activities: Net Cash Provided By Used In Operating Activities [Abstract] Net loss Profit Loss Adjustments to reconcile net loss to net cash used in operating activities: Adjustments To Reconcile Net Income Loss To Cash Provided By Used In Operating Activities [Abstract] Depreciation Depreciation Equity-based compensation expense Share Based Compensation Accretion/amortization of investment securities Accretion Amortization Of Discounts And Premiums Investments Reduction in carrying amount of operating lease right of use asset Reduction In Carrying Amount Of Operating Lease Right Of Use Asset Changes in operating assets and liabilities: Increase Decrease In Operating Capital [Abstract] Accounts receivable Increase Decrease In Accounts Receivable Prepaid expenses and other current assets Increase Decrease In Prepaid Deferred Expense And Other Assets Prepaid research and development, net of current portion Increase Decrease In Prepaid Research And Development Net Of Current Portion Accounts payable and accrued expenses Increase Decrease In Accounts Payable And Accrued Liabilities Deferred revenue Increase Decrease In Contract With Customer Liability Operating lease liabilities Increase Decrease In Operating Lease Liability Other long-term liabilities Increase Decrease In Other Noncurrent Liabilities Other assets Increase Decrease In Other Operating Assets Net cash used in operating activities Net Cash Provided By Used In Operating Activities Cash flows from investing activities: Net Cash Provided By Used In Investing Activities [Abstract] Purchases of marketable securities Payments To Acquire Available For Sale Securities Debt Proceeds from maturity of marketable securities Proceeds From Sale And Maturity Of Marketable Securities Proceeds from redemption of marketable securities Proceeds From Maturities Prepayments And Calls Of Available For Sale Securities Purchases of property and equipment Payments To Acquire Property Plant And Equipment Net cash (used in) provided by investing activities Net Cash Provided By Used In Investing Activities Cash flows from financing activities: Net Cash Provided By Used In Financing Activities [Abstract] Payments on finance lease obligations Finance Lease Principal Payments Proceeds from issuance of common stock, net of issuance costs Proceeds From Issuance Of Common Stock Proceeds from employee stock purchases and exercise of stock options Proceeds From Employee Stock Purchases And Exercise Of Stock Options Payment of employee withholding taxes relating to restricted stock awards Payments Related To Tax Withholding For Share Based Compensation Net cash provided by financing activities Net Cash Provided By Used In Financing Activities Net increase in cash, cash equivalents and restricted cash Cash Cash Equivalents Restricted Cash And Restricted Cash Equivalents Period Increase Decrease Including Exchange Rate Effect Cash, cash equivalents and restricted cash at beginning of period Cash Cash Equivalents Restricted Cash And Restricted Cash Equivalents Including Disposal Group And Discontinued Operations Cash, cash equivalents and restricted cash at end of period Supplemental disclosure of non-cash investing activities: Noncash Investing Items [Abstract] Assets acquired under operating lease obligation Assets Acquired Under Operating Lease Obligation Property and equipment purchases included in accounts payable and accrued expenses Noncash Purchase Of Property Plant And Equipment In Accounts Payable And Accrued Liabilities Adjustments Supplemental disclosure of non-cash financing activities: Noncash Financing Items [Abstract] Purchase under finance lease Capital Lease Obligations Incurred Issuance costs included in accounts payable and accrued expenses Issuance Costs Included In Accounts Payable And Accrued Expenses Cash paid for interest Interest Paid Net Organization Consolidation And Presentation Of Financial Statements [Abstract] Nature of Business Organization Consolidation And Presentation Of Financial Statements Disclosure [Text Block] Accounting Policies [Abstract] Summary of Significant Accounting Policies Significant Accounting Policies [Text Block] Fair Value Disclosures [Abstract] Fair Value of Financial Instruments Fair Value Disclosures [Text Block] Available for sale securities disclosure. Investments Debt And Equity Securities [Abstract] Available-for-Sale Investments Available For Sale Securities Disclosure [Text Block] Property Plant And Equipment [Abstract] Property and Equipment, Net Property Plant And Equipment Disclosure [Text Block] Payables And Accruals [Abstract] Accrued Expenses Accounts Payable Accrued Liabilities And Other Liabilities Disclosure Current [Text Block] Equity [Abstract] Stockholders' Equity Stockholders Equity Note Disclosure [Text Block] Disclosure Of Compensation Related Costs Sharebased Payments [Abstract] Equity-based Compensation and Equity Incentive Plans Disclosure Of Compensation Related Costs Share Based Payments [Text Block] Collaboration Agreements Collaborative Arrangement Disclosure [Text Block] Earnings Per Share [Abstract] Net Loss per Share Earnings Per Share [Text Block] Commitments And Contingencies Disclosure [Abstract] Commitments and Contingencies Commitments And Contingencies Disclosure [Text Block] Related Party Transactions [Abstract] Related-Party Transactions Related Party Transactions Disclosure [Text Block] Basis of Presentation Basis Of Accounting Policy Policy [Text Block] Principles of Consolidation Consolidation Policy [Text Block] Recently adopted accounting pronouncements. Recently Adopted Accounting Pronouncements Recently Adopted Accounting Pronouncements Policy [Text Block] Recently Issued Pronouncements New Accounting Pronouncements Policy Policy [Text Block] Schedule of Company's Classified Assets Measured at Fair Value on Recurring Basis Schedule Of Fair Value Assets And Liabilities Measured On Recurring Basis Table [Text Block] Summary of Available-for-Sale Securities Held Schedule Of Available For Sale Securities Reconciliation Table [Text Block] Schedule of Property and Equipment, Net Property Plant And Equipment [Text Block] Schedule of Accrued Expenses Schedule Of Accrued Liabilities Table [Text Block] Schedule of shares of common stock reserved for future issuance. Schedule of Shares of Common Stock Reserved for Future Issuance Schedule Of Shares Of Common Stock Reserved For Future Issuance Table [Text Block] Schedule of Stock Option Activity Schedule Of Share Based Compensation Stock Options Activity Table [Text Block] Schedule of Restricted Common Stock Activity Nonvested Restricted Stock Shares Activity Table [Text Block] Schedule of Equity-based Compensation Expenses Schedule Of Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Text Block] Schedule of Equity-based Compensation Expenses by Award Type Disclosure Of Share Based Compensation Arrangements By Share Based Payment Award [Text Block] Schedule of Potentially Common Shares Excluded from Calculation of Net Loss Per share Schedule Of Antidilutive Securities Excluded From Computation Of Earnings Per Share [Text Block] Cash, cash equivalents, and short-term marketable securities Cash Cash Equivalents And Short Term Investments Accumulated deficit Summary of significant accounting policies. Summary of significant accounting policies. Summary Of Significant Accounting Policies [Table] Summary Of Significant Accounting Policies [Table] Accounting Standards Update Adjustments For New Accounting Pronouncements [Axis] Accounting Standards Update Type Of Adoption [Member] ASU 2017-08 Accounting Standards Update201708 [Member] Summary Of Significant Accounting Policies [Line Items] Summary Of Significant Accounting Policies [Line Items] Change in accounting principle, accounting standards update, adopted [true false] Change In Accounting Principle Accounting Standards Update Adopted Change in accounting principle, accounting standards update, adoption date Change In Accounting Principle Accounting Standards Update Adoption Date Change in accounting principle, accounting standards update, immaterial effect [true false] Change In Accounting Principle Accounting Standards Update Immaterial Effect Fair Value Asset Measured On Recurring Basis [Line Items] Fair Value Asset Measured On Recurring Basis [Table] Fair Value Asset Measured On Recurring Basis [Table] Fair Value Asset Measured On Recurring Basis [Table] Measurement Frequency Fair Value By Measurement Frequency [Axis] Measurement Frequency Fair Value Measurement Frequency [Domain] Recurring Fair Value Measurements Recurring [Member] Fair Value Hierarchy and NAV Fair Value By Fair Value Hierarchy Level [Axis] Fair Value Hierarchy and NAV Fair Value Measurements Fair Value Hierarchy [Domain] Quoted Prices in Active Markets for Identical Assets (Level 1) Fair Value Inputs Level1 [Member] Significant Other Observable Inputs (Level 2) Fair Value Inputs Level2 [Member] Significant Unobservable Inputs (Level 3) Fair Value Inputs Level3 [Member] Financial Instrument Financial Instrument [Axis] Financial Instruments Transfers And Servicing Of Financial Instruments Types Of Financial Instruments [Domain] Corporate Debt Securities Corporate Debt Securities [Member] Investment Type Investment Type [Axis] Investments Investment Type Categorization [Member] Commercial Paper Commercial Paper [Member] US treasury and government long-term debt securities. U.S. Government Agency Securities and Treasuries U S Treasury And Government Long Term Debt Securities [Member] Cash and Cash Equivalents Cash And Cash Equivalents [Axis] Cash and Cash Equivalents Restricted Cash And Cash Equivalents Cash And Cash Equivalents [Member] Money market funds Money Market Funds [Member] Fair Value Asset Measured On Recurring Basis [Line Items] Fair Value Asset Measured On Recurring Basis [Line Items] Cash and Cash Equivalents Cash And Cash Equivalents Fair Value Disclosure Marketable securities Marketable Securities Short-term investments Investments Fair Value Disclosure Total assets Assets Fair Value Disclosure Debt Securities Available For Sale [Table] Debt Securities Available For Sale [Table] U.S. government agency securities U S Government Agencies Debt Securities [Member] Schedule of Available-for-sale Securities [Line Items] Schedule Of Available For Sale Securities [Line Items] Amortized cost Available For Sale Debt Securities Amortized Cost Basis Gross unrealized gains Available For Sale Debt Securities Accumulated Gross Unrealized Gain Before Tax Gross unrealized losses Available For Sale Debt Securities Accumulated Gross Unrealized Loss Before Tax Fair Value Available For Sale Securities Debt Securities Other than temporary impairment losses investments number debt securities available for sale. Number of investments in unrealized loss position Debt Securities Available For Sale Unrealized Loss Position Number Of Positions Number of investments in unrealized loss position, more than twelve months Debt Securities Available For Sale Continuous Unrealized Loss Position12 Months Or Longer Number Of Positions Aggregate fair value of securities in unrealized loss position Debt Securities Available For Sale Unrealized Loss Position Number of securities with other than temporary impairment Other Than Temporary Impairment Losses Investments Number Debt Securities Available For Sale Schedule Of Property Plant And Equipment [Table] Schedule Of Property Plant And Equipment [Table] Long-Lived Tangible Asset Property Plant And Equipment By Type [Axis] Long-Lived Tangible Asset Property Plant And Equipment Type [Domain] Laboratory Equipment Equipment [Member] Computer and office equipment. Computer and Office Equipment Computer And Office Equipment [Member] Furniture and Fixtures Furniture And Fixtures [Member] Leasehold Improvements Leasehold Improvements [Member] Construction in Progress Construction In Progress [Member] Property Plant And Equipment [Line Items] Property Plant And Equipment [Line Items] Property and equipment, gross Property Plant And Equipment Gross Less accumulated depreciation Accumulated Depreciation Depletion And Amortization Property Plant And Equipment Property and equipment, net Accrued research and development, current. Payroll related Employee Related Liabilities Current Professional fees Accrued Professional Fees Current Research and development Accrued Research And Development Current Other Other Accrued Liabilities Current Total accrued expenses Class of warrant or right exercise price of warrants paid at closing of offering. Proceeds from issuance of common stock and pre funded warrants. Warrants exercised. Schedule Of Stock By Class [Table] Schedule Of Stock By Class [Table] Related Party Related Party Transactions By Related Party [Axis] Related Party Related Party [Domain] Ginkgo Bioworks, Inc. Ginkgo Bioworks, Inc. Ginkgo Bioworks Inc [Member] Class of Warrant or Right Class Of Warrant Or Right [Axis] Class of Warrant or Right Class Of Warrant Or Right [Domain] Pre-Funded Warrants. Pre-Funded Warrants Pre Funded Warrants [Member] Common Stock at the market. ATM At The Market [Member] Cowen and company LLC. Cowen and Company, LLC Cowen And Company L L C [Member] Jefferies, LLC Jefferies, LLC Jefferies L L C [Member] Underwritten public offering. Underwritten Public Offering Underwritten Public Offering [Member] Class Of Stock [Line Items] Class Of Stock [Line Items] Sale of common stock, shares Sale of stock, price per share Sale Of Stock Price Per Share Warrants to purchase shares of common stock Class Of Warrant Or Right Number Of Securities Called By Warrants Or Rights Warrants exercise price per share Class Of Warrant Or Right Exercise Price Of Warrants Or Rights1 Warrants exercise price per share paid at closing of offering Class Of Warrant Or Right Exercise Price Of Warrants Paid At Closing Of Offering Proceeds from issuance of common stock and pre-funded warrants Proceeds From Issuance Of Common Stock And Pre Funded Warrants Warrants exercised Warrants Exercised Common Stock Issuable Under Pre-Funded Warrants Option Indexed to Issuer's Equity Option Indexed To Issuers Equity Equity [Axis] Option Indexed to Issuer's Equity, Type Option Indexed To Issuers Equity Type [Domain] Options exercisable to purchase common stock. Options Exercisable to Purchase Common Stock Options Exercisable To Purchase Common Stock [Member] Employee Stock Purchase Plan Employee Stock [Member] Total Shares of Common Stock Reserved for Future Issuance Common Stock Capital Shares Reserved For Future Issuance Schedule Of Share Based Compensation Arrangements By Share Based Payment Award [Table] Schedule Of Share Based Compensation Arrangements By Share Based Payment Award [Table] Plan Name Plan Name [Axis] Plan Name Plan Name [Domain] Long Term Incentive Plan 2015. Plan 2015 Long Term Incentive Plan2015 [Member] ESPP Two thousand seventeen and two thousand fifteen equity incentive award plan. 2017 Plan and 2015 Plan Two Thousand Seventeen And Two Thousand Fifteen Equity Incentive Award Plan [Member] Award Type Award Type [Axis] Award Type Share Based Compensation Arrangements By Share Based Payment Award Award Type And Plan Name [Domain] Restricted Common Stock Restricted Stock [Member] Share-based Compensation Arrangement by Share-based Payment Award [Line Items] Share Based Compensation Arrangement By Share Based Payment Award [Line Items] Increase in number of shares available for issuance (in shares) Share Based Compensation Arrangement By Share Based Payment Award Number Of Additional Shares Authorized Options exercised to purchase shares of common stock under ESPP Shares available for future grants Share Based Compensation Arrangement By Share Based Payment Award Number Of Shares Available For Grant Outstanding Beginning balance, Number of options Share Based Compensation Arrangement By Share Based Payment Award Options Outstanding Number Granted, Number of options Share Based Compensation Arrangement By Share Based Payment Award Options Grants In Period Gross Exercised, Number of options Cancelled/Forfeited, Number of options Share Based Compensation Arrangement By Share Based Payment Award Options Forfeitures In Period Outstanding Ending balance, Number of options Number of options, Vested or expected to vest Share Based Compensation Arrangement By Share Based Payment Award Options Vested And Expected To Vest Outstanding Number Number of options, Exercisable Share Based Compensation Arrangement By Share Based Payment Award Options Exercisable Number Beginning balance, Weighted-average price Share Based Compensation Arrangement By Share Based Payment Award Options Outstanding Weighted Average Exercise Price Granted, Weighted-average price Share Based Compensation Arrangements By Share Based Payment Award Options Grants In Period Weighted Average Exercise Price Exercised, Weighted-average price Share Based Compensation Arrangements By Share Based Payment Award Options Exercises In Period Weighted Average Exercise Price Cancelled/Forfeited, Weighted-average price Share Based Compensation Arrangements By Share Based Payment Award Options Forfeitures In Period Weighted Average Exercise Price Ending balance, Weighted-average price Weighted-average price Vested or expected to vest Share Based Compensation Arrangement By Share Based Payment Award Options Vested And Expected To Vest Outstanding Weighted Average Exercise Price Weighted-average price, Exercisable Share Based Compensation Arrangement By Share Based Payment Award Options Exercisable Weighted Average Exercise Price Outstanding, weighted average remaining contractual term (Year) Sharebased Compensation Arrangement By Sharebased Payment Award Options Outstanding Weighted Average Remaining Contractual Term2 Weighted average remaining contractual term, Vested or expected to vest Sharebased Compensation Arrangement By Sharebased Payment Award Options Vested And Expected To Vest Outstanding Weighted Average Remaining Contractual Term1 Weighted average remaining contractual term, Exercisable Sharebased Compensation Arrangement By Sharebased Payment Award Options Exercisable Weighted Average Remaining Contractual Term1 Share based compensation arrangement by share based payment award options forfeitures and expirations in period intrinsic value. Beginning balance, Aggregate Intrinsic value Share Based Compensation Arrangement By Share Based Payment Award Options Outstanding Intrinsic Value Exercised, Aggregate Intrinsic value Share Based Compensation Arrangement By Share Based Payment Award Options Exercises In Period Total Intrinsic Value Cancelled/Forfeited, Aggregate Intrinsic value Share Based Compensation Arrangement By Share Based Payment Award Options Forfeitures And Expirations In Period Intrinsic Value Ending balance, Aggregate Intrinsic value Aggregate Intrinsic value, Vested or expected to vest at September 30, 2021 Share Based Compensation Arrangement By Share Based Payment Award Options Vested And Expected To Vest Outstanding Aggregate Intrinsic Value Aggregate Intrinsic value, Exercisable at September 30, 2021 Sharebased Compensation Arrangement By Sharebased Payment Award Options Exercisable Intrinsic Value1 Share based compensation by share based arrangement award, options available for aggregate intrinsic value. Share-based compensation arrangement by share-based payment award, options outstanding, weighted average exercise price, and additional disclosures. Share Based Compensation Arrangement By Share Based Payment Award Options Outstanding Weighted Average Exercise Price And Additional Disclosures [Abstract] Number of options in money Share Based Compensation By Share Based Arrangement Award Options Available For Aggregate Intrinsic Value Employee unrecognized compensation expense Employee Service Share Based Compensation Nonvested Awards Total Compensation Cost Not Yet Recognized Stock Options Employee unrecognized compensation cost, period of recognition Employee Service Share Based Compensation Nonvested Awards Total Compensation Cost Not Yet Recognized Period For Recognition1 Beginning balance, Number of unvested shares/units Share Based Compensation Arrangement By Share Based Payment Award Equity Instruments Other Than Options Nonvested Number Granted, Number of shares/units Share Based Compensation Arrangement By Share Based Payment Award Equity Instruments Other Than Options Grants In Period Vested, Number of shares/units Share Based Compensation Arrangement By Share Based Payment Award Equity Instruments Other Than Options Vested In Period Forfeited, Number of shares/units Share Based Compensation Arrangement By Share Based Payment Award Equity Instruments Other Than Options Forfeited In Period Ending balance, Number of unvested shares/units Beginning balance, Unvested Grant date fair value Share Based Compensation Arrangement By Share Based Payment Award Equity Instruments Other Than Options Nonvested Weighted Average Grant Date Fair Value Granted, Grant date fair value Share Based Compensation Arrangement By Share Based Payment Award Equity Instruments Other Than Options Grants In Period Weighted Average Grant Date Fair Value Vested, Grant date fair value Share Based Compensation Arrangement By Share Based Payment Award Equity Instruments Other Than Options Vested In Period Weighted Average Grant Date Fair Value Forfeited, Grant date fair value Share Based Compensation Arrangement By Share Based Payment Award Equity Instruments Other Than Options Forfeitures Weighted Average Grant Date Fair Value Ending balance, Unvested Grant date fair value Schedule Of Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Table] Schedule Of Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Table] Income Statement Location Income Statement Location [Axis] Income Statement Location Income Statement Location [Domain] Research and Development Expense Research And Development Expense [Member] General and Administrative Expense General And Administrative Expense [Member] Share-based Compensation Arrangement by Share-based Payment Award, Compensation Cost [Line Items] Employee Service Share Based Compensation Allocation Of Recognized Period Costs [Line Items] Equity-based compensation expense Allocated Share Based Compensation Expense Stock Options Employee Stock Option [Member] Restricted Stock Awards Collaboration agreement upfront, nonrefundable technology access fee. Potential milestones payment upon achievement of certain success criteria. Schedule Of Collaborative Arrangements And Noncollaborative Arrangement Transactions [Table] Schedule Of Collaborative Arrangements And Noncollaborative Arrangement Transactions [Table] Collaborative Arrangement and Arrangement Other than Collaborative Type Of Arrangement [Axis] Collaborative Arrangement and Arrangement Other than Collaborative Arrangements And Nonarrangement Transactions [Member] Roche collaboration and option agreement. Roche Collaboration and Option Agreement Roche Collaboration And Option Agreement [Member] Statistical Measurement Range [Axis] Statistical Measurement Range [Member] Minimum Minimum [Member] Maximum Maximum [Member] Balance Sheet Location Balance Sheet Location [Axis] Balance Sheet Location Balance Sheet Location [Domain] Current liabilities. Current Liabilities Current Liabilities [Member] Ginkgo Collaboration Abb Vie agreement. AbbVie Agreement Abb Vie Agreement [Member] Business Acquisition Business Acquisition [Axis] Business Acquisition, Acquiree Business Acquisition Acquiree [Domain] IBD Programs. IBD Programs I B D Programs [Member] Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items] Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items] Upfront, nonrefundable technology access fee Collaboration Agreement Upfront Nonrefundable Technology Access Fee Milestones payment upon achievement of certain success criteria Potential Milestones Payment Upon Achievement Of Certain Success Criteria Anticipated completion period for research and development projects. Collaboration in research and development Anticipated Completion Of Research And Development Projects Period Upfront payments. Upfront non refundable payment Upfront Payments Milestone Payments Milestone payments Milestone Payments Recognition of revenue Current deferred revenue Prepayment to related party for collaboration agreement. Related party transaction collaboration agreement, initial term. Prepayment to related party for collaboration agreement Prepayment To Related Party For Collaboration Agreement Related party transaction collaboration agreement, initial term Related Party Transaction Collaboration Agreement Initial Term Current pre-paid research and development Prepaid Expense Current Non-current pre-paid research and development Prepaid Expense Noncurrent Number of shares sold Schedule Of Antidilutive Securities Excluded From Computation Of Earnings Per Share [Table] Schedule Of Antidilutive Securities Excluded From Computation Of Earnings Per Share [Table] Antidilutive Securities Antidilutive Securities Excluded From Computation Of Earnings Per Share By Antidilutive Securities [Axis] Antidilutive Securities, Name Antidilutive Securities Name [Domain] Unvested Restricted Stock Awards [Member] Unvested Restricted Common Stock Awards Unvested Restricted Stock Awards [Member] Employee stocks purchase plan. Potential Shares Issuable Under the ESPP Employee Stocks Purchase Plan [Member] Antidilutive Securities Excluded From Computation Of Earnings Per Share [Line Items] Antidilutive Securities Excluded From Computation Of Earnings Per Share [Line Items] Potentially Common Shares Excluded from Calculation of Net Loss Per share Antidilutive Securities Excluded From Computation Of Earnings Per Share Amount Schedule Of Related Party Transactions By Related Party [Table] Schedule Of Related Party Transactions By Related Party [Table] Related Party Transaction [Line Items] Related Party Transaction [Line Items] Current Pre-Paid Research and Development. Non Current Pre-Paid Research and Development Prepaid research and development expenses. Prepayment to related party for collaboration agreement Related party transaction collaboration agreement, initial term Current Pre-Paid Research and Development Current Pre Paid Research And Development Non Current Pre-Paid Research and Development Non Current Pre Paid Research And Development Prepaid research and development expenses Prepaid Research And Development Expenses